Plague decimates the crowded city, where many have arrived to embark for the New World to seek their fortune. Mateo Nunez is asked to discharge a debt to a dead friend - to bring his illegitimate son from the city. As Mateo seeks out the boy, Celso de Guevara is taken through the overcrowded slums and shown the body of a murdered man whose marked hands are holding a crucifix.

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Mateo is brought before the grand inquisitor and asked to find the murderer of one of an important trader.

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The body of a murdered woman is found in the harbour of Seville. The woman turns out to be Lucina Utrera - the daughter of a sea captain who is away with the fleet on his way home from America. Mateo discovers that the two murders are linked.

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The plague devastates the city. Mateo and Valerio find the missing lover of Ignacio Azuaga, Carmen Galvez. She confirms to them that Ledesma - the Protestants's secret leader - seems to be behind all the murders.

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Mateo and Valerio exhume German's corpse and corroborate he was also murdered by Ledesma. Zuniga takes advantage of the plague, the famine and the popular uprisings to negotiate with the Assistant and persuade him to open the city's harbour to allow the fleet from America to unload the shipments.

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As the epidemic starts to decline, Inquisitor General Celso de Guevara hands Mateo the bull granting his freedom, but the case is far from over.

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